Asp.net MVC + routelink只得到404错误

下面是我的代码:

RouteConfig.cs

public class RouteConfig
{
    public static void RegisterRoutes(RouteCollection routes)
    {
        routes.IgnoreRoute("{resource}.axd/{*pathInfo}");
        routes.MapRoute(
            name: "snakes",
            url: "snakes-databases/",
            defaults: new { controller = "SnakesController", action = "index" }
        );
        routes.MapRoute(
            name: "snakesLists",
            url: "snakes-databases/{family}/",
            defaults: new { controller = "SnakesController", action = "Lists", family = "" }
        );
        routes.MapRoute(
            name: "snakesDetails",
            url: "snakes-databases/{family}/{species}/",
            defaults: new { controller = "SnakesController", action = "Details", family = "", species = "" }
        );
        routes.MapRoute(
            name: "Default",
            url: "",
            defaults: new { controller = "Home", action = "Index" }
        );
    }
}

_Layout.cshtml

@Html.RouteLink("snakes database", "snakes", null, new { @class = "dropdown-toggle", data_toggle="dropdown"})
<ul class="dropdown-menu">
    <li>@Html.RouteLink("Boidae", "snakesLists", new { family = "boidae" })</li>
    <li>@Html.RouteLink("Colubridae", "snakesLists", new { family = "colubridae" })</li>
    <li>@Html.RouteLink("Elapidae", "snakesLists", new { family = "elapidae" })</li>
    <li>@Html.RouteLink("Viperidae", "snakesLists", new { family = "viperidae" })</li>
</ul>

SnakeController.cs:

public ActionResult Lists(string family)
{
    return View();
}

我不明白为什么每个url除了主页(/)我得到一个404错误。